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Phyllocephalinae

CLASSIFICATION OF THE PHYLLOCEPHALINAE

There has been very little work on the higher classification of this subfamily until recent times. Linnavuori (1982) presented an extensive discussion of the phylogeny of African genera, but refrained from using actual tribal names. Kamaluddin and Ahmad in a series of papers have erected several tribal names to contain the Indian genera. Their classification does not totally agree with that of Linnavuori, but between the two, with some intuition, most phyllocephaline genera can be placed to tribe.

Linnavuori placed the African genera in three generic groups, but indicated that there were still three genera that seemed to represent their own independent phylogenetic lineage. At the time of Linnavuoriís work only one tribal name was available, Delocephalini. Actually Delocephalini was erected by HorvŠth in 1900 as a tribe in the family Tessaratomidae, but Kumar (1969) transferred the type genus, Delocephalus, to the Phyllocephalinae of the Pentatomidae without commenting on the tribal situation. Linnavuoriís groupings were:

Phyllocephala Group
Lamtoplax
Linnavuori
Melampodius
Distant
Nimboplax
Linnavuori
Phyllocephala
Laporte

Lobopeltista Group
Dichelorhinus
St
Dl
Frisimelica
Distant
Gellia
St
Dl
Gonopsis
Amyot & Serville
Katongoplax
Linnavuori
Lobopeltista
Schouteden
Macrina
Amyot & Serville
Sandehana
Distant

 

Dalsira Group [= Delocephalini ?]
Chalcopis
Kirkaldy
Dalsira
Amyot & Serville [= Basicryptus]
Delocephalus
Distant
Eonymia
Linnavuori
Melanocryptus
Linnavuori [= Jayma]
Metocryptus
Linnavuori
Metonymia
Kirkaldy [= Dalsira]
Penedalsira
Linnavuori
Schismatops
Dallas
Tantia
Distant
Tshibalaka
Schouteden

Independent Lineages
Kafubu
Schouteden
Schyzops
Spinola
Storthogaster
Karsch

 

Ahmad and his associates, in a series of papers worked extensively on the higher classification of the subfamily, erecting three new tribal names, and assigned a number of genera to the nominate tribe. Their classification is:

Cressonini Kamaluddin & Ahmad
Cressona
Dallas
Kafuba
Schouteden
Melampodius
Distant
Nimboplax
Linnavuori

Megarrhamphini Ahmad & Kamaluddin
Bakerorandolotus
Ahmad & Kamaluddin
Megarrhamphus
Bergroth
Randolotus
Distant

 

Phyllocephalini Amyot & Serville
Dalsira
Amyot & Serville
Diplorhinus
Amyot & Serville
Gonopsis
Amyot & Serville
Mercatus
Distant
Metonymia
Kirkaldy
Nazeeriana
Ahmad & Kamaluddin
Salvianus
Distant
Schyzops
Spinola

Tetrodini Ahmad & Kamaluddin
Gellia
Tetroda
Tetrodias
Kirkaldy

 

Basically, Ahmadís classification goes something like this. Those genera which have the humeral angles greatly expanded and produced anteriorly with the anterolateral margins toothed or crenulate belong in the Cressonini; the very elongate slender species belong in the Megarrhamphini; the species which have the lateral margins of the head expanded and laminate belong in the Tetrodini; and all others would belong in the Phyllocephalini. By combining the above two classifications with a little intuition, we have arrived at the following classification for this catalog:

Cressonini Kamaluddin & Ahmad
Cressona
Dallas
Kafuba
Schouteden
Lamtoplax
Linnavuori
Melampodius
Distant
Nimboplax
Linnavuori

Megarrhamphini Ahmad & Kamaluddin
Bakerorandolotus
Ahmad & Kamaluddin
Megarrhamphus
Bergroth
Randolotus
Distant

Tetrodini Ahmad & Kamaluddin
Gellia
St
Dl
Tetroda
Tetrodias
Kirkaldy

 

Phyllocephalini Amyot & Serville
Basicryptus
Herrich-Sch
šffer
Borrichias
Distant
Chalcopis
Kirkaldy
Dalsira
Amyot & Serville
Delocephalus
Distant
Dichelorhinus
St
Dl
Diplorhinus
Amyot & Serville
Eonymia
Linnavuori
Frisimelica
Distant
Gonopsimorpha
Yang
Gonopsis
Amyot & Serville
Jayma
Rider
Melanocryptus
Linnavuori
Kaffraria
Kirkaldy
Magwamba
Distant
Katongoplax
Linnavuori
Lobopeltista
Schouteden
Macrina
Amyot & Serville
Mercatus
Distant
Metocryptus
Linnavuori
Metonymia
Kirkaldy [= Dalsira]
Minchamia
Gross
Nazeeriana
Ahmad & Kamaluddin
Neoschyzops
Ahmad & Kamaluddin
Penedalsira
Linnavuori
Phyllocephala
Laporte
Roebournea
Schouteden
Salvianus
Distant
Sandehana
Distant
Schismatops
Dallas
Schyzops
Spinola
Storthogaster
Karsch
Tantia
Distant
Tshibalaka
Schouteden
Uddmania
Bergroth

 

 

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David A. Rider
Professor of Entomology
North Dakota State University
202 Hultz Hall
Fargo, ND 58105
E-Mail: David.Rider@ndsu.edu

updated: 30 Aug 2005 

Published by the Department of Entomology 


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